Sound ?
Lot of question marks here.. how intelligent are these toe-ticklers.. They live like wolves in packs, so it's rather an animal. Maybe it likes the sound ? just the right high pitch frequency ? By
Chimpansees like the waygiggling sound, because it is a sign of "non-attack", refer article link below.
I don't think this could be a truely wild hunter.. although humans and apes and rats appreciate tickling, I doubt effective tickling could happen without consent ! What's the role of toe-ticklers in human society ? How come these animals are allowed to tickle girls.. The toe-tickler could be the girl's pet.. or ismaybe it is some kind of cultural habit humans learned, because wild ticklerstoe-ticklers are really good at tickling toes ? But in that case why only girls and not the rest of us mammals, who like ticklingbeing tickled and make high pitch sounds like e.g. rats do, or my grandmother does (I'm lost now)
http://www.bbc.com/earth/story/20170518-why-humans-chimpanzees-and-rats-enjoy-being-tickled