The only things that I know about this kind of animals is that comes from animals that originally were not social and was convergent in many different animals, so till where I know the polimorphic variations of the castes are caused for the same reasons which produce "races", for a specie with a high genetical variability in this case brought to the extreme and combined with posterior polifenism changes caused by the enviroment.
Then I could concluse that for get this eusocial "elemental" creature is required start with a specie in which just the individual variations produce the very different "breathings". But I am nor sure if this breathing structures can be developed after the "eusocialization" as a subproduct of this.
So, maybe the primordial structures that the queen should have for produce the other castes can be limited to just three, because according to what I read they can be used as precursors to others because changing or adding some things can have multiple functions. These are:
- Electric, because can be used as lighter for develop lightning, fire or napalm.
- Napalm or other substance, because have the "container" for create the required substances for be throwing.
- Cryogenic, is the mores strange, but probably change the orientiation, can chane from "ice" to "heat" or to air currents.
Of curse how I mentioned these structures can be developed after the eusocialization but for this case I do not have options.
Finally for mention the ideas for the castes:
- The bigger soldiers being fire breather.
- The workers, could be ice or cryogenic breathers for build some structures (but maybe this need a big "refrigerator" body), the other option a sticky substance like napalm could work.
- Light drones, maybe would be electric or even ligthning throwers.
- And I already mentioned the options for the queen.
Many other characteristics can be added but that could push even more the limits of the polimorphism, so we will be atenied to a very similar body format with just size varitions but with this different "breathings".
Also there are a lot of more dragons or thrower animals that I did not want include because if I dont the principals less I know that, but I am gonna let those here for reference.